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All rights reserved/ ECOSPHERE 11(9) :e03251(2020); doi:10.1002/ecs2.3251 ©2020 The Authors. This is an open access article under the terms of the Creative Commons AttributionLicense, which permits use, distribution and reproduction in any medium, provided the original work is properly cited./ BMC ECOLOGY AND EVOLUTION 21(1) :64(2021); doi:10.1186/s12862-021-01796-8 © The Author(s) 2021. Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. 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